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various points in the game, the player is able to choose between good and evil paths which take a fair amount of time to complete. There are multiple endings depending on how the game is played out. In addition, the player can choose which villains to fight against. The player is able to switch between Spider-Man's traditional red and blue costume and the symbiote suit at any time during gameplay.[4] Chris Scholz from Shaba Games also commented that the developers "worked on really separating the black and red suits to give the player a different feel." For example, red and blue-suited Spider-Man is fast and agile, while symbiote Spider-Man is stronger and more destructive. The mini-map on the screen is also different from the style used in previous Spider-Man games. The traditional 2-D overhead view has been removed, replaced by a new 3D map with an "X", "Y" and "Z" axis which shows the player's position in three-dimensional space (north and south, as well as vertical position). Also, as a Wii exclusive, there are 6 alternate costumes (Spider-Carnage for the black suit, and Spider-Armor, Ben Reilly, Spider-Man 2099, Cosmic Spider-Man and the Iron Spider suit for the red suit).[5]Spider-Man searches for Mary Jane Watson while the city of New York is infested with symbiotes. He finds her with Luke Cage but then sees a mysterious figure and is attacked. During a flashback to four days earlier, Spider-Man retells his account of how his latest encounter with Venom ended with a piece of the symbiote binding to himself. After the fight with Venom, an injured Mary Jane is sent to a hospital and Spidey meets Luke Cage during a gang war. Cage explains the Rolling Sevens and Park Avenue gangs are at war, and that their task is to bring the two to a truce. After pursuing the two gang leaders across the city, Spider-Man manages to get them to agree to a peaceful meeting in a park in Harlem.

Outside the rendezvous point, Spider-Man discovers high-tech assassins on top of the nearby buildings. After defeating them, he traces the assassins back to the Kingpin. He ends up running into the Black Cat, who appears to be working with the Kingpin. The two fight briefly, but in the end, Black Cat reveals she is working against Kingpin. The two join forces with Moon Knight in order to stop the Kingpin's illegal activities, battling several of his men, including the Vulture. Eventually, Spider-Man confronts a number of Kingpin's henchman on the roof of the Daily Bugle, where a number of civilians suddenly attack them with their hands and teeth. A news helicopter sees Spider-Man throwing civilians off of the building, and the police believe him to be a murderer. Unknown to them, the civilians have been infected with symbiotes.

Spider-Man web-slinging through a symbiote-infested New York City. As the game progresses, symbiote constructs will appear

Wolverine, thinking Spider-Man to be possessed by a symbiote, tracks down and confronts the web-slinger in Hell's Kitchen. After a battle, the two settle their differences and begin tracking down infected citizens. Eventually, Spider-Man finds Venom throwing people into symbiote pods, turning them into monsters. The two engage in a cross-city battle in which Venom escapes and several of the pods are destroyed. A S.H.I.E.L.D. team led by Black Widow arrives quarantines New York and sets up symbiote controlled citizen quarantine camps. Electro attacks one of the camps looking for his sister, who accidentally infects him with the symbiote attached to her.

Since no genius heroes are available, Spider-Man decides to break the Tinkerer out of Riker's to utilise his scientific genius to find a countermeasure to the symbiotes. Upon reaching the prison, Spider-Man teams up with the Rhino, using his strength to help free the Tinkerer. The Tinkerer (in return for a pardon, etc.) then designs a device to be installed on the Trask Building; it will not harm the infected hosts, only remove the symbiotes. While the Tinkerer constructs the device, Spider-Man aids S.H.I.E.L.D. in dealing with the symbiote infestation in the streets of Manhattan. First, he confronts the symbiote Electro in Central Park, defeating him and stopping him from reaching the city's power supply. He then helps Wolverine in evacuating a church filled with civilians, but Wolverine ends up getting infected and attacks Spider-Man. Spider-Man is able to purge Wolverine of the symbiote, and then goes looking for Mary Jane

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